Tortoise Species

Plasma Steroid and Nutrient Levels in Wild Testudo Horsfieldi
2003 article on seasonal variations in sex steroids, proteins etc. as measured in Horsfields in the wild.

Food habits of the Leopard Tortoise
Geochelone Pardalis observed in Tanzania.

Manouria Emys
Observations on the Asian Brown tortoise observed in the wild.

Eastern Box Turtle
Some observations on the North American Eastern Box Turtle (Terapene carolina carolina), including thoughts on decline of various local populations.

Foraging behaviour and diet of an ectothermic herbivore: Testudo horsfieldi
An excellent article on the eating habits and diet of the Russian tortoise. ECOGRAPHY 26: 236–242, 2003

Proper Diet for Desert Tortoises
Excellent and broad-ranging article that goes beyond the diet of the Desert Tortoise to cover many of the principles of good tortoise diets. By Geoff Stein (palmbob)

Effect of Supplemental Heat in Captive African Leopard Tortoises (Stigmochelys Pardalis) & Spurred Tortoises (Centrochelys Sulcata) on Growth Rate & Carapacial Scute Pyramiding
A research article which concludes that increased heat exposure during the night contributes directly to pyramiding of the carapacial scutes and that when caring for tortoises in captive environments, attention should be paid to the use of variable day/night temperature heat & cooling cycles. (Mark L. Heinrich, MS, DVM, and Kaleb K. Heinrich, MS, DA Dec. 2015)

The Egyptian Tortoise, Testudo kleinmanni Lortet, 1883 in Libya
This article publishes the observations of a research team studying the population of Egyptian (Testudo Kleinmanni) in Libya and includes a list of plants eaten and plants used for shelter, as well as an assessment of the most potent threats to this species in the wild. (Christoph Schneider & Willi Schneider, 2008)

Seasonal changes in the diet of Testudo Hermanni Hermanni in central Italy
An interesting observational study of wild Hermanni Hermanni, including seasonal differences in diet and gender-specific preferences.

Hermanni Haven, Garden State Tortoise
An excellent comprehensive site dedicated to all things Hermanni, including guidance on various Hermanni subspecies, as well as other species in the genus Testudo. (Chris Leone, Garden State Tortoise)

Desert Tortoises (Gopherus agassizii) Are Selective Herbivores that Track the Flowering Phenology of Their Preferred Food Plants
Foraging Habits of the Desert Tortoise. Observational study of the diet of wild Desert Tortoises (Gopherus agassizii) in the western Mojave Desert, USA.